BULLETIN DE LTNSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE

BULLETIN VAN HET KONINKLIJK BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN

SCIENCES DE LA TERRE AARDWETENSCHAPPEN VOL. 72

BRUXELLES 2002 BRUSSEL

BULLETIN DE L’INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE

BULLETIN VAN HET KONINKLIJK BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN

SCIENCES DE LA TERRE AARDWETENSCHAPPEN VOL. 72

BRUXELLES 2002 BRUSSEL

> TUM

R édacteur en ch e f - H oofdredacteur - Editor. A nnie V . D h o n d t

Secrétaire de rédaction - R edactiesecretaris - A ssocia te editor: Jacques G o d e f r o i d

C om ité de rédaction - R edactiecom ité - E ditoria l board : Pierre B u l t y n c k

Daniel C a h e n

M ichel D f.l ie n s

C om ité in ternational - Internationaal com ité - C onsulting ed ito rs : A leksandr S. A l e k s e e v ( M o s c o w , Russia)

D enise B r ic e (L ille, France)Jenaro L. GARCiA-ALCADE(Oviedo, Spain)

M ichael A. K a m i n s k i (London, UK)Jordi M a r t i n e l l (B arcelona, Spain)

D avid B. W e i s h a m p e l (B altim ore, U SA )

La rédaction rem ercie pour lecture critique de m anuscrits:De redactie dankt volgende review ers voor m edew erking aan dit volum e:

For this volum e the fo llow ing review ers are gratefully acknow ledged:

R. B. B l o d g e t t , D. B o h a s k a , D e n is e B r ic e , C.H .C . B r u n t o n , P. C a r l s , Jackie C i a n o , R. J. C l e e v e l y ,

G . J. D y k e , R. F e i s t , R. F e l d m a n n , O . H a m p e , R. K a e s t l e r , E.V. K u r o c h k i n , B. M i s t i a e n ,

C écile M o u r e r - C h a u v i r é , C. N e u m a n n , W .A. O l iv e r J r ., S . L. O l s o n , X . P e r e d a S u b e r b i o l a , A.B. S m i t h ,

D. T s h u d y , Tatiana A. T u m a n o v a

BU LLETINDE L ’IN STITU T RO Y A L DES SC IEN C ES N A T U R E L L E S DE B ELG IQ U E

SC IEN C ES DE LA TER R E

BU LLETINVAN HET K O N IN K LIJK B ELG ISCH IN STITU U T V O O R N A T U U R W E T E N SC H A PPE N

A A R D W ETEN SC H A PPEN

Vol. 72 - 2002 ISSN 0374-6291

Publié, verschenen, p u b lish ed : 3 LU I.2002

© E dition de l ' Institut Royal des Sciences N aturelles

de B elgique Rue V autier 29

B -1000 B ruxelles, Belgique

© U itgave van het K oninklijk B elgisch Instituut voor

N atuurw etenschappen V autierstraat 29

B-1000 Brussel, België

TABLE DES MATIÈRES CONTENTS

A v e r i a n o v , A . O. - A n an k y lo sau rid (O m ith i- schia: A nky losau ria) b ra incase from the U pper C re taceous B issek ty F orm ation o f U zbek istan 97

C a s i e r , J .G ., L e t h i e r s , F. & P r é a t , A . -

O straco d s and sed im en to lo g y o f the D ev o ­n ian -C arb o n ife ro u s s tra to ty p e sec tion (L a S erre , M o n tag n e N o ire , F rance) 43

C o e n - A u b e r t , M . - T en in o p h y llid s and Spi- n o p h y llid s (R u g o sa) from the G iv e tian M ont d ’H aurs F o rm atio n in B elg ium 5

G o d e f r o i d , J., S t a i n i e r , P. & T r o s t , G. - T w o new sp in o cy rtiid b rach io p o d s (S p irife ri- da) from the P rag ian and the L o w er E m sian o f B elg ium 25

J a g t , J .W .M . - P erip rocta l p la tin g in the ech i- no id C o en h o lec typ u s lo r te ti (C en o m an ian , S u ltan a te o f O m an) 85

J a g t , J.W .M . & F r a a i j e , R .H .B . - T he erym id lobster E nop lo c ly tia lea ch ii ( M a n t e l l , 1822) from the U p p er C am p an ian o f no rtheast B e l­g ium 91

K a z â r , E. - R ev ised p h y lo g en y o f the P h y se ­te rid ae (M am m alia : C e tacea ) in the ligh t o f P la co zip h iu s V a n B e n e d e n , 1869 and A ulo - p h y se te r K e l l o g g , 1927 151

K o w a l k e , T. - S y stem atic rev is io n o f P alaeo- cen e b rack ish w a te r G astro p o d a from M ons, B elg ium , b ased on th e ir early on to g en e tic shells 111

M a y r , G. & S m i t h , R. - A new reco rd o f the P ro p h ae th o n tid ae (A ves: P e lecan ifo rm es)from the M idd le E ocene o f B elg iu m 135

M a y r , G. & S m i t h , R. - A v ian rem ain s from the low erm o st O lig o cen e o f H oogbu tse l (B e l­g iu m ) 139

S a r t e n a e r , P. & P l o d o w s k i , G . - D esc rip ­tion o f M eso ch o risp ira ko n in c k i ( D e w a l - q u e , 1895), a sp irife rid b rach io p o d from the B elg ian U p p e r T o u m a is ian 69

V e r h e y d e n , T. - D ecapods from the B oom C lay (R u p e lian , O lig o cen e) in B elg iu m 171

BULLETIN DE L'INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE BULLETIN VAN HET KONINKLIJK BELGISCH INSTITUUT VOOR NATUURW ETENSCHAPPEN

SCIENCES DE LA TERRE, 72: 5-24, 2002 AARDW ETENSCHAPPEN, 72: 5-24, 2002

Temnophyllids and Spinophyllids (Rugosa) from the Givetian Mont cTHaurs Formation in Belgium

by M arie C O E N -A U B E R T

C o e n - A u b e r t , M., 2002. - T em nophyllids and Spinophyllids (Rugosa) from the G ivetian M ont d 'H au rs Form ation in B elgium . Bulletin de l 'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre, 72: 5-24, 3 pis., 6 figs., Bruxelles-B russel, M arch 31, 2002. - ISSN 0374-6291.

A b s t r a c t

Temnophyllum majus W a l t h e r , 1 9 2 9 , T. imperfectum n. sp., Spino- phyllum spongiosum ( S c h l ü t e r , 1 8 8 9 ) and S. blacourti ( R o h a r t , 1 9 8 8 ) are described in detail and com e m ainly from the low er part o f the G ivetian M ont d ’Haurs Form ation, on the south side o f the Dinant Synclinorium . T he type specim ens o f S. spongiosum, Temnophyllum majus, T. latum W a l t h e r , 1 9 2 9 and T. clavatum W a l t h e r , 1 9 2 9 as w ell as the holotype o f Spinophyllum altevogti ( O l i v e r & S o r a u f , 1 9 8 8 ) are refigured herein. A s a w hole, the fauna from the M ont d ’H aurs Form ation, w hich belongs to the Low er Polygnathus varcus Zone, is w idely d istributed in various areas o f Europe and A sia and show s strong affin ities w ith the G ivetian rugose corals o f G erm any and the B oulonnais in France.

Key-words: R ugose corals, G ivetian, Taxonom y, Stratigraphy, Bel­gium.

Résumé

Temnophyllum majus W a l t h e r , 1 9 2 9 , T. imperfectum n. sp., Spino­phyllum spongiosum ( S c h l ü t e r , 1 8 8 9 ) et 5 . blacourti ( R o h a r t , 1 9 8 8 ) sont décrits en détail et proviennent principalem ent de la partie infé­rieure de la Form ation givetienne du M ont d 'H aurs , au bord sud du Synclinorium de D inant. Les spécim ens types de 5. spongiosum, Temnophyllum majus, T. latum W a l t h e r , 1 9 2 9 et T. clavatum W a l t h e r , 1 9 2 9 de m êm e que l ’holotype de Spinophyllum altevogti ( O l i v e r & S o r a u f , 1 9 8 8 ) sont refigurés dans ce travail. L ’ensem ble de la faune de la Form ation du M ont d ’H aurs, qui appartient à la Z one à Polygnathus varcus inférieure, est largem ent répandue dans diverses régions d ’Europe et d ’A sie et m ontre de nettes affin ités avec les Rugueux givetiens d ’A llem agne et du Boulonnais en France.

M o ts-c le fs : Rugueux, G ivetien, T axinom ie, S tratigraphie, Belgique.

Introduction

After the papers o f C o e n - A u b e r t (1 9 9 9 and 2 0 0 0 ) mainly devoted to colonial rugose corals, it is interesting to investigate the rich fauna o f solitary rugose corals occurring in the Mont d ’Haurs Formation from the south side o f the Dinant Synclinorium. The following taxa are

d e s c r ib e d h e re in : Temnophyllum majus W a l t h e r , 1 9 2 9 , T. imperfectum n . sp ., Spinophyllum spongiosum (S C H L Ü ­

T E R , 1 8 8 9 ) a n d 5. blacourti (R o h a r t , 1 9 8 8 ). S. spongio­sum is th e ty p e s p e c ie s o f Spinophyllum W e d e k i n d , 1922 w h e r e a s Temnophyllum majus is c o m p a r e d to th e o th e r s p e c ie s o f Temnophyllum W a l t h e r , 1 9 2 9 in t r o d u c e d by W a l t h e r ( 1 9 2 9 ) .

From a stratigraphical point o f view, the Mont d ’Haurs Formation belongs to the Lower Polygnathus varcus conodont Zone and lies in the upper part o f the Givetian between the Terres d ’Haurs and From elennes Formations (Fig. 6). From a geographical point o f view and as it was the case before, most specimens come from the area between Beauraing and Han-sur-Lesse located to the east o f Givet (Fig. 1). However, some coralla have been collected in three poorly known outcrops o f Nismes situated 5 km to the northeast o f Couvin. M oreover, the north side o f the Dinant Synclinorium and the Nam ur Synclinorium characterized by different facies have also provided a few samples.

The main part o f the material was collected by the author in situ during geological surveys made bed by bed. This sampling is supplemented by old thin sections re­ferred in this paper to the “ Old collection from the Institut royal des Sciences naturelles de Belgique” . The types o f the new species and the figured specimens are also stored in the collections o f the Institut royal des Sciences naturelles de Belgique (IRScNB).

D e s c r i p t i o n o f t h e o u t c r o p s

N o r t h e r n l e s l i m i t e s q u a r r y a t A v e -e t -A u f f e

(W ellin M C -1988-6; Fig. 2)

This active quarry is the most com plete section investi­gated in the M ont d ’Haurs Formation. It has been located on a map and described previously by C o e n - A u b e r t

(1999, p. 27, figs. 2, 3 and 2000, p. 5, fig. 2).At the top o f the Terres d ’Haurs Formation, which

consists o f coarsely crinoidal limestone, there are already some rugose corals represented by Sociophyllum wede- kindi C o e n - A u b e r t , 1999, Acanthophyllum simplex

6 Marie COEN-AUBERT

( W a l t h e r , 1929) and Spinophyllum spongiosum. Then, the Mont d ’Haurs Formation reaches a thickness o f about 180 m and is subdivided into two parts.

The limestone is rather argillaceous and often very rich in corals and strom atoporoids in the lower part o f the lithostratigraphic unit which is 91 m thick. At the top o f these 91 m, there is a double level o f thin-bedded lime­stone which has a thickness o f 10 m and which is a key bed in the area between Beauraing and Han-sur-Lesse. The rugose coral fauna from the lower part o f the Mont d 'H aurs Formation exhibits a great diversity and shows some vertical distribution. S. spongiosum is restricted to the first 37 m o f the lithostratigraphic unit whereas Dis- phyllum semenoffi C o e n - A u b e r t , 2000 has only been found between 16 m and 28 m above its base. Other species such as S. blacourti, Acanthophyllum simplex and Sociophyllum wedekindi occur throughout the lower part o f the Mont d ’Haurs Formation. S. isactis ( F r e c h ,

1886), Temnophyllum majus and T. imperfectum have also been observed up to the second level o f thin-bedded limestone, but appear respectively 17 m, 22 m and 35 m above the top o f the Terres d ’Haurs Formation. Argutas- trea tenuiseptata C o e n - A u b e r t & L ü t t e , 1990 is very common in the upper 45 m from the lower part o f the Mont d ’Haurs Formation where it is locally accompanied by A. wangi ( T s i e n , 1978). Grypophyllum denckmanni W e d e k i n d , 1922 is rather rare in the northern Les Limites quarry.

In the upper part o f the Mont d ’Haurs Formation which is 89.5 m thick, the rugose corals are less abundant and are

represented by Argutastrea wangi, Wapitiphyllum laxum ( G ü r ic h , 1896) and Sunophyllum beichuanense H e , 1978.

W e l l in q u a r r y (W ellin M C -1986-4; Fig 2)

The disused quarry o f W ellin has been located on a map by C o e n - A u b e r t (1999, fig. 2) and described in detail by C o e n - A u b e r t (2000, p. 6, fig. 2). Below the first level o f thin-bedded limestone from the Mont d ’Haurs Formation are exposed 40 m o f argillaceous and rather pure lime­stone which contain a highly diversified fauna o f corals and stromatoporoids. Disphyllum sem enoffi is restricted to the base o f this sequence. Acanthophyllum sim plex , Grypophyllum denckmanni and Sociophyllum wedekindi are observed throughout these 40 m. S. isactis, Temno­phyllum m ajus, T. imperfectum and Spinophyllum bla­courti are very abundant in the middle part o f these reefal limestones whereas Argutastrea tenuiseptata is only pre­sent in their upper 11 m.

Above the first level o f thin-bedded limestone, there is a lack o f outcrop which is 28.5 m thick. In its middle however, one can see 0.85 m o f argillaceous or dolomitic limestone which contains massive stromatoporoids, a few corallites o f Sociophyllum isactis and S. wedekindi and numerous coralla o f Temnophyllum majus. This bed be­longs already to the upper part o f the Mont d ’Haurs Formation. At the end o f the section, there are about 10 m o f biostromal limestone with intercalations o f fine limestone, where a few specimens o f T. majus and Gry­pophyllum denckmanni still occur.

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BRABANT MASSIFV E R V I E R S ?

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Fig. 1 — G en era l s ituation in the sou th o f B elgium .

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Fig. 2 Comparative logs ot the Beauraing quarry, the Wellin quarry, the northern Les Limites quarry at Ave-et-Auffe and the Han-sur-Lesse section with the distribution of rugose corals. (For explanation of conventional signs, see Fig. 3).

Mont d’H

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8 Marie COEN-AUBERT

C O N V E N T I O N A L S I G N S

S T R O M A T O P O R O I D S O CRINOIDS

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Fig. 3 — Explanation of conventional signs used in Figs. 2 and 5.

B e a u r a in g q u a r r y (Beauraing M C-1975-3; Fig. 2)

The Mont d ’Haurs Formation is exposed in the second quarry excavated to the south o f Beauraing, along the road to Winenne. The outcrop has been located on a map and described by C o e n - A u b e r t (1999, p. 29, figs. 3 and 5).

Below the first level o f thin-bedded limestone, there is a discontinuous section showing 24 m o f rather argillac­eous limestone, very rich in corals and stromatoporoids. Argutastrea tenuiseptata, Sociophyllum isactis, S. wede­kindi and rare Grypophyllum denckmanni occur in this part o f the sequence. One corallum o f Spinophyllum blacourti has been collected at the base o f the thin-bedded limestones whereas Argutastrea tenuiseptata and Temno­phyllum imperfectum are present between these two le­vels. Then, the upper part o f the Mont d ’Haurs Formation starts with about 10 m o f dolomitic limestone which contains numerous massive and dendroid strom atopor­oids, alveolitids, tham noporids and solitary rugose corals including T. imperfectum and T. majus.

H a n -s u r - L e s s e s e c t io n (Han-sur-Lesse M C -1975-15; Fig. 2)

The Han-sur-Lesse section in the M ont d ’Haurs Forma­tion is situated along the road to Rochefort and has also been located on a map and described by C o e n - A u b e r t

(1999, p. 29, figs. 3 and 6). Sociophyllum isactis has been observed at the base o f the outcrop which is very dis­

continuous. In the 30 m o f reefal and often argillaceous limestones, which lie below the key layers o f thin-bedded limestones, occur S. isactis, S. wedekindi, Temnophyllum imperfectum and Argutastrea tenuiseptata accompanied locally by A. wangi. A. tenuiseptata and Temnophyllum imperfectum are present with rare Grypophyllum denck­manni in the coralliferous beds between the two levels o f thin-bedded limestones. A few specimens o f Temnophyl­lum imperfectum, Acanthophyllum sim plex and Socio­phyllum wedekindi have been collected up to 20 m above the base from the upper part o f the Mont d 'H aurs Forma­tion.

R e s t e ig n e q u a r r y (W ellin M C -1974-95)

In the disused quarry o f Resteigne, the base o f the Mont d ’Haurs Formation is exposed in continuity with the Terres d ’Haurs Formation. This transition has been de­scribed and figured by C o e n - A u b e r t (1999, p. 29 and 2000, fig. 2). As it is the case in the northern Les Limites quarry, Spinophyllum spongiosum and Acanthophyllum simplex are present at the top o f the Terres d ’Haurs Formation which is also characterized by coarsely crinoi- dal limestone. C o e n - A u b e r t (2000) has mentioned at the base o f the M ont d ’Haurs Formation the occurrence o f A. simplex, Grypophyllum denckmanni and Sociophyllum wedekindi.

A r e a o f N i s m e s (Figs. 4 and 5)

The three outcrops investigated in the Mont d ’Haurs Formation at Nismes have been shown to the author by Coen and Dumoulin who are revising the geological map o f Olloy-sur-Viroin - Treignes.

Along the west and east sides o f the big excavation called Matricolo (outcrop Olloy-sur-Viroin MC-65) and lying in the Bois Mousti, to the southwest o f Nismes, there are two good sections in the upper 42 m from the lower part o f the Mont d ’Haurs Formation. At the top o f this sequence, some thin-bedded limestones occur, but this level is not so remarkably developed as in the area between Beauraing and Han-sur-Lesse. Below this key level, the

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Fig. 4 — Location of the three outcrops investigated at Nis­mes.

Mont d ’Haurs Formation 9

MONT D’HAURS FORMATION AT NISMES

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Fig. 5 — Comparative logs of the three outcrops investigated at Nismes with the distribution of rugose corals. (For explanation of conventional signs, see Fig. 3).

more or less argillaceous limestones are rich in corals and stromatoporoids and contain Argutastrea tenuiseptata, So­ciophyllum isactis, S. wedekindi, Acanthophyllum simplex, Temnophyllum majus and T. imperfectum.

A similar succession is exposed in the small hole (out­crop Olloy-sur-Viroin MC-64 and point Olloy 466 o f Coen and Dumoulin) located 1250 m to the northeast o f Nismes. Just above the equivalent o f the thin-bedded limestones appear crinoidal limestones with reef building organisms in some layers and two bedding planes full o f solitary rugose corals including T. imperfectum and T. majus.

In another excavation (outcrop Olloy-sur-Viroin MC- 63 and point Olloy 12 from the old collection o f the IRScNB) lying 400 m to the east o f Nismes, one can see the top o f the reefal and argillaceous limestones from the lower part o f the M ont d ’Haurs Formation overlain by3.5 m o f more or less thin-bedded limestones. After a lack o f outcrop which is about 7 m thick, the upper part o f the M ont d 'H aurs Formation is exposed with a thickness o f 66 m and is characterized by an alternation o f fine or bioclastic limestones and biostromes with massive stro­matoporoids accompanied by dendroid stromatoporoids, thamnoporids, scolioporids and solitary rugose corals.

These coralla are not very well preserved in this outcrop whereas colonies o f Argutastrea tenuiseptata and A. wangi are present near the top o f the lower part from the M ont d ’Haurs Formation.

N o r t h s i d e o f t h e D in a n t s y n c l in o r iu m a n d N a m u r

SY N CLIN O R IU M

Several specimens have also been collected at the local­ities o f Cour-sur-Heure, Gerpinnes and Aisemont inves­tigated by C o e n - A u b e r t ( 2 0 0 0 , pp. 8 -1 1 , figs. 4 -6 ) . Though Cour-sur-Heure belongs to the northern part o f the Dinant Synclinorium, its Givetian facies are sim ilar to those from the south side o f the same synclinorium. In the southern quarry o f Cour-sur-Heure (outcrop Gozée MC- 47), Spinophyllum blacourti is present in the Terres d ’Haurs Formation, 13 m below its top whereas Temno­phyllum imperfectum has been found close to the top from the lower part o f the M ont d ’Haurs Formation, in the vicinity o f the conodont sample mentioned by C o e n - A u b e r t ( 2 0 0 0 , p. 9 ).

Gerpinnes is situated 11 km to the northeast o f Cour-sur- Heure, also on the north side o f the Dinant Synclinorium.

10 Marie COEN-AUBERT

However, the Givetian deposits are very different here as the Mont d ’Haurs Formation passes laterally into the Névre- mont Formation. At the base o f the disused railway section from Gerpinnes (outcrop Nalinnes MC-1975-2), T. imper­

fectum and Spinophyllum blacourti occur together with Acanthophyllum simplex and Argutastrea tenuiseptata, in 16 m o f argillaceous and more or less reefal limestones by which the outcrop starts in the Névremont Formation.

Along the railway section at Aisemont (outcrop Ta- mines M C -1983-2), on the south side o f the Nam ur Synclinorium, there are, close to the base o f the N évre­mont Formation, 12 m o f argillaceous limestones and shales which contain brachiopods, crinoids and several beds rich in rugose corals represented by A. tenuiseptata, Acanthophyllum simplex, Grypophyllum denckmanni, Spinophyllum spongiosum, S. blacourti and Temnophyl­lum imperfectum.

Finally, one corallum o f T. imperfectum has been sampled at the depth o f 1242 m, in the borehole o f Feuze ( 125E/298) described by C o e n - A u b e r t et al. ( 1981 ). The

city o f Leuze lies in the Tertiary from the western part o f Belgium, 16 km to the east o f Tournai; its palaeozoic basem ent belongs to the north side o f the N am ur Syncli- norium. In fact, T. imperfectum has been observed in the Alvaux Formation which is the lateral equivalent o f the Névremont Formation in this area. In the Leuze borehole, the Alvaux Formation has been intersected between 1157 m and 1373 m and is 216 m thick. It is characterized by an alternation o f argillaceous limestones and shales with a few beds o f anhydrite. Besides T. imperfectum, Argutastrea tenuiseptata has been recognized at 1363 m, near the base o f the lithostratigraphic unit.

S t r a t i g r a p h i e d i s t r i b u t i o n o f t h e r u g o s e c o r a l s

On the south side o f the Dinant Synclinorium, the lower part o f the Mont d ’Haurs Formation is characterized by a highly diversified fauna o f rugose corals (Fig. 6). Some species such as Sociophyllum wedekindi, Acanthophyllum

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Mont d ’Haurs Formation 11

simplex and Spinophyllum spongiosum are already pre­sent at the top o f the underlying Terres d ’Haurs Form a­tion. Then, S. spongiosum and Disphyllum semenoffi are restricted to the base o f the Mont d ’Haurs Formation whereas Sociophyllum wedekindi, Acanthophyllum sim ­plex, Grypophyllum denckmanni and Spinophyllum bla­courti are observed throughout the lower part o f the lithostratigraphic unit. Other taxa such as Sociophyllum isactis, Temnophyllum majus and T. imperfectum are also very common in the lower part o f the M ont d ’Haurs Formation, but do not occur at its base. Argutastrea tenuiseptata appears rather high in the sequence and is locally accompanied by A. wangi which is more frequent in the upper part o f the Mont d ’Haurs Formation. At the base o f these deposits, the fasciculate and solitary rugose corals remain rather abundant and are still represented by several species from the lower part o f the lithostrati­graphic unit. Higher in the Mont d 'H aurs Formation, the fauna o f the biostromes intercalated with back reef facies is dominated by massive stromatoporoids. How­ever, at the top o f the lithostratigraphic unit, the rugose and tabulate corals become again numerous with the occurrence o f Wapitiphyllum laxum and Sunophyllum beichuanense.

On the north side o f the Dinant Synclinorium and on the south side o f the Nam ur Synclinorium, the Mont d ’Haurs Formation passes laterally into the Névremont Formation. Acanthophyllum simplex, Grypophyllum denckmanni, Temnophyllum imperfectum, Spinophyllum spongiosum, S. blacourti and Argutastrea tenuiseptata are reported close to its base whereas A. wangi and Wapitiphyllum laxum characterize its upper part.

B i o g e o g r a p h i c i m p l i c a t i o n s

The rugose corals investigated in the Mont d ’Haurs For­mation by C o e n - A u b e r t (1999 and 2000) and herein show strong affinities with the Givetian faunas from the Boulonnais in France and from the Eifel Hills, the Ber- gisches Land and the Sauerland in Germany. Firstly, it is to be mentioned that five species have been introduced in Germany, namely Sociophyllum isactis, Temnophyllum majus, Spinophyllum spongiosum, Acanthophyllum sim ­plex and Grypophyllum denckmanni. In the Eifel Hills, the three latter taxa and Argutastrea tenuiseptata, whose stratigraphie distribution has been discussed by C o e n - A u b e r t & L ü t t e (1990, p. 23), are known in the Rodert and Kerpen Formations or at least in one o f them. In the Bergisch Gladbach-Paffrath syncline from the Bergisches Land, A. tenuiseptata is recorded in the Torringen For­mation whereas Grypophyllum denckmanni, Spinophyl­lum spongiosum and Sociophyllum isactis have been collected in the Büchel Formation. About S. isactis, it must be noted that its lectotype figured by W e d e k in d (1925, pi. 4, fig 12) has been found again by B ir e n h e id e

(1998, p. 179). Unfortunately, this information remained unknown to C o e n - A u b e r t (1999, p. 34) before the pub­lication o f her paper though the material o f the Ardennes

was compared with that o f W e d e k i n d (1925). In the North Bergisches Land, Acanthophyllum sim plex is re­ported in the Schwelm Formation and at the top o f the Givetian by B ir e n h e id e (1990). Also in the Schwelm Formation, but in the North Sauerland occur A. simplex, Grypophyllum denckmanni, Temnophyllum majus and Spinophyllum spongiosum. On the other hand and in the same area, the colony referred to Argutastrea tenuisepta­ta by M a y (1993, p. 28) and sampled near the Eifelian- Givetian boundary differs from the Belgian specimens by rather thick septa. Finally, it is well known that the Rodert and Torringen Formations are roughly time-equivalent to the Terres d ’Haurs Formation whereas the Kerpen, Bü­chel and Schwelm Formations are more or less o f the same age as the Mont d 'H aurs Formation. These data are confirmed by the recent Devonian Correlation Table published by W e d d ig e (1996: R160dm96, 1998: R002dm97 and R012dm97 and 2000: R410dm00).

Besides Spinophyllum blacourti defined in this area, Acanthophyllum simplex, Wapitiphyllum laxum , Argutas­trea wangi and A. tenuiseptata are present in the Blacourt Formation from the Boulonnais. According to C o e n - A u ­

b e r t & L ü t t e (1990, p. 22), the latter taxon was de­scribed as Disphyllia periclada ( K r ä m e r , 1982) by R o -

HA RT (1988). Moreover, the stratigraphie distribution o f these rugose corals, as stated by R o h a r t (1988, tab 2) in the Griset M ember o f the Blacourt Formation, is rather sim ilar to that observed in the Mont d ’Haurs Formation from Belgium.

In the Givetian o f Great Britain, Acanthophyllum sim ­p lex is reported in W est Somerset whereas Temnophyllum majus and Grypophyllum denckmanni have been recog­nized in South Devon. The latter taxon is also mentioned by Jo s e p h & T s ie n (1975, p. 190) in the Givetian o f the Pyrenees in France. To the south o f Europe, Acantho­phyllum simplex has been collected by C o e n - A u b e r t (2002, p. 34) in the Upper Givetian o f the Tafilalt and the M a’der in Morocco.

In Poland, Wapitiphyllum laxum has been introduced in the Upper Givetian o f Dziewki from Silesia. According to W r z o l e k (1993, p. 232), it is associated with Temno­phyllum majus identified by the author as 7'. latum W a l t h e r , 1929. Both species were also sampled by W r z o l e k (1993) in the Upper Givetian o f the Holy Cross Mountains. Furthermore, Wapitiphyllum laxum is highly characteristic o f the Upper Givetian from M oravia in the Czech Republic, as mentioned by G a l l e (1985, p. 55). Sociophyllum isactis was observed by K e t t n e r o v a (1932, p. 47) in the Givetian o f the same area whereas Grypophyllum denckmanni has been found by G a l l e

(1994, p. 45) in the Acanthopyge limestone o f Bohemia, also in the Czech Republic. This lithostratigraphic unit is traditionally assigned to the Eifelian, but it is possible that it belongs to the Givetian after the last data o f conodonts given by G a l l e (1994, p. 42).

Farther to the east, Sunophyllum beichuanense has been defined in the Givetian o f the Sichuan Province in China whereas Sociophyllum isactis and Grypophyllum denckmanni have a wide geographic range according to

12 Marie COEN-AUBERT

C o e n - A u b e r t ( 1 9 9 9 , p. 3 3 a n d 2 0 0 0 , p . 14). In d e e d , o n e o r b o th o f th e s e s p e c ie s occur:

in the Givetian o f the Russian Platform and the Urals in Russia;in the Givetian o f the Tien Shan, Guizhou and Yunnan in China as well as in the M iddle Devonian o f North­east China and the Gansu Province.Moreover, Sociophyllum isactis has been reported in

the Givetian o f New South W ales and North Queensland in Australia. So it appears that many taxa identified in the Mont d ’Haurs Formation from Belgium are present in various areas from the Old World Realm and occasionally in Australia whereas none o f them is recorded in the Middle Devonian o f North America.

S y s t e m a t i c P a l a e o n t o l o g y

Family D is p iiy l l id a e H il l , 1939 Genus Temnophyllum W a l t h e r , 19 2 9

Type speciesBy subsequent designation o f L a n g et al. (1940, p. 132), Temnophyllum latum W a l t h e r , 1929.

D ia g n o s is

Solitary rugose corals. Septa o f two orders, non-carinate or sometimes faintly carínate. Both orders o f septa fre­quently in lateral contact in the outer part o f the dissepi- m entarium so as to form a wide peripheral stereozone which is com plete or partial. Septa more or less dilated in the inner part o f the dissepimentarium and thin in the tabularium. M ajor septa reaching the axis o f the corallum or leaving an open space in the centre o f the tabularium. Minor septa traversing the entire dissepimentarium. Dis­sepim entarium composed o f several rows o f globose dis­sepiments, often arranged in horizontal layers in its outer part and inclined towards the axis o f the corallum in its inner part. Tabulae usually incomplete.

Tem nophyllum m ajus W a l t h e r , 1929 Plate 1, Figures 3-13, Plate 2, Figures 13, 14

v * 1929 Temnophyllum major- W a l t h e r , p. 127.V 1929 Temnophyllum clavatum- W a l t h e r , p . 128,

fig. 18.v 1929 Temnophyllum nodosum- W a l t h e r , p . 125,

figs. 15, 16.v non 1929 Temnophyllum latum- W a lth er , p. 123, fig . 14.v 1975 Temnophyllum latum- T s ie n , fig. 16a.v p. 1977 Temnophyllum latum- T s ie n , figs. 4o, p (non

fig. 4n = Temnophyllum imperfectum n. sp. herein ).

v 1978 Temnophyllum majus Walther 1928- B ir e n ­h e id e , p . 8 6 , p i. 12, fig. 2.

p. 1985 Temnophyllum majus Walther 1928- B ir e n ­h e id e & L ia o , p. 242 (non pi. 3, fig. 13).

LectotypePI. 12, figs. 2b, c in B ir e n h e id e (1978) and pi. 1, figs. 3 ,4

figured herein, chosen by B ir e n h e id e & L ia o (1985, p. 242). Specimen and thin sections SMF W DKD 6995- 6997 stored in the Forschungsinstitut Senckenberg at Frankfurt am Main, Germany. Upper Givetian Schwelm Formation at Sundwig near Iserlohn in North Sauerland, Germany.

Material and localitiesEighty specimens with 124 thin sections. Personal sampling: Wellin MC-1988-6-A719, A721, A723, A725. B4. BX7 and B 114; Wellin MC-1986-4-A508B, A508C, A508L, A510D, A510E, A510F, A526H, A532D, A532J, A532P, A532Q, A532T, A532U, A532V, A535D, A544A, A544B, A544C, A546, A5841, A5842, A5843, A5846, A58410, A58411, A58412, A58416. A58417 and A58423; Beauraing MC-1975- 3-Z703, Z706 and Z707; Han-sur-Lesse MC-1975-15-Z943; Olloy-sur-Viroin MC-64-C839; Olloy-sur-Viroin MC-65- C867 and C896. Old collection from the Institut royal des Sciences naturelles de Belgique: Agimont (Mont d’Haurs at Givet)-Gi-28566, 28580, 28584, 28589, 28591, 28601, 28603, 28606, 28609, 28668, 28694, 28959, 28963, 28966, 28967, 28969, 28970, 28974, 28978, 28981, 28982, 28985, 28993, 28995 and 29004; Olloy-sur-Viroin 12-Gid-11894, 11896, 11915, 11917, 11923, 11929, 11935, 11953, 11959, 12012, 12163 and 12368.

D ia g n o s is

A species o f Temnophyllum with 56 to 66 septa at a diameter o f 11 mm to 20 mm. Wide and continuous stereozone at the periphery forming a ring against the wall and obscuring the outer part o f the dissepimentarium.

D e s c r ip t io n

The material consists o f com plete or fragmentary coralla which are often cylindrical and sometimes conical, tochoid or ceratoid. Their height varies between 1.5 cm and 4.5 cm, but may reach 7 cm. Longitudinal ribs have been observed in a few specimens and one o f them shows constrictions due to rejuvenescence. The outer wall is not always well preserved, but is frequently en­crusted by thin laminar stromatoporoids or rarely by alveolitids.

A wide and continuous stereozone, where the septa are contiguous laterally, occurs systematically in the outer part o f the dissepim entarium though some voids appear locally at the periphery o f several coralla. Beyond the stereozone, the septa are non-carinate and dilated in the dissepim entarium and become thinner in the tabularium. Occasionally, they are dilated throughout their length, less thick in the tabularium or again dilated at their axial ends. The m ajor septa reach the axis o f the corallum or leave a more or less extensive open space in the centre o f the tabularium. In very few specimens, one can observe a weak whorl, pseudofossulae, a plane o f bilateral sym m e­try, trabeculae or isolated fragments o f septa. The minor septa traverse the entire dissepim entarium or even enter into the tabularium where they may be contratingent; they are rarely shorter.

The dissepim entarium consists o f 4 to 14 rows o f small globose dissepim ents which are in horizontal layers at the periphery and inclined in its inner part. Its outer part is

Mont d 'Haurs Formation 13

normally obscured by contiguous coarse trabeculae which are subhorizontal or slightly inclined towards the outer wall. The tabulae are incomplete and intersecting laterally; sometimes their general pattern is concave or convex.

There are 52 to 70 septa per corallum. The diam eter o f the corallum ranges from 8 mm to 26 mm. The width o f the tabularium varies commonly between 5.5 mm and 10 mm and more generally between 4.5 mm and 11.5 mm.

D i s c u s s io n

Up to now, Temnophyllum majus was only known by two specimens from the type locality o f Sundwig in the Sauer- land, Germany (PI. 1, figs. 3 -6 ) . According to B i r e n h e i d e

(1 9 7 8 , p . 8 6 ) , it differs only by larger coralla from T. latum o f the Givetian Oberhonsel or Schwelm Formation from the same area and type species o f Temnophyllum. But the holotype o f T. latum (PI. 1, figs. 1, 2) figured among others by B i r e n h e i d e & L i a o ( 1 9 8 5 , pi. 7, fig. 3 8 ) shows clearly that the peripheral stereozone is not con­tinuous in transverse section as in T. majus. This particu­larity has been described in detail by P i c k e t t ( 19 6 7 , p. 6 7 ) and has been observed by the writer. However, several identifications o f T. latum in the literature seem to con­cent coralla o f T. majus. This is for instance the case for the material o f M i d d l e t o n (1 9 5 9 . p. 1 5 4 ) which comes from the Givetian o f South Devon in Great Britain and for the material o f W r z o l e k ( 1 9 9 3 , p. 2 3 2 ) which comes from the Givetian o f Silesia and the Holy Cross Moun­tains in Poland and maybe from the Upper Frasnian o f the latter arca. This is also the case for the two transverse sections o f T. latum illustrated by T s i e n (1 9 7 5 and 1 9 7 7 ) and collected in the Mont d 'H aurs Formation at Givet in France.

The diameters o f the Belgian specimens referred to T. majus are often not so large as those o f the lectotype and paralectotype. They are more similar to those o f the coralla o f T. clavatum W a l t h e r , 1929 (PI. 1, fig. 7) and T. nodosum W a l t h e r , 1929 figured by W a l t h e r

( 1929) and also recorded in the Givetian o f the Sauerland. In these two taxa, the wide peripheral stereozone is complete as in T. majus. On the other hand, this stereo­zone is poorly developed in the sample assigned to the latter species by B ir e n h e id e & L ia o (1985) and found in the Givetian o f the Guizhou Province in China.

T. altum ( W a l t h e r , 1929) from the Lower Frasnian o f North Sauerland, that has been revised by B ir e n h e id e & L ia o (1985, p. 240), is closely related to T. majus. How­ever, its outer stereozone invests nearly the entire disse­pimentarium so that the minor septa project only locally beyond it. Moreover, it is characterized by slightly smal­ler size and septal number and by axial tabcllae system­atically convex.

A more or less complete stereozone is developed in the outer part o f the dissepimentarium o f several other taxa. In Entelophyllum alpinum V o n S c h o u p p é (1951) from the Givetian o f the Camic Alps in Austria, that probably belongs to the genus Temnophyllum and that was de­scribed by V o n S c h o u p p é (1954, p. 403), the septa are

not typically contiguous in the wide stereozone. Addition­ally, they are less numerous in this species which is only represented by its not well preserved holotype. In the two specimens o f Temnophyllum etheridgei Z iie n & J e l l ,

1996 from the Givetian o f North Queensland in Australia illustrated by Z h e n «fe J e l l (1996, pi. 12, figs. 7 and 8), the peripheral stereozone is partially destroyed by weath­ering; these two coralla are also rather small with slightly fewer septa. In Tortophyllum zhongguoense J ia , 1977 from the Devonian o f the Hunan Province in China in­vestigated by J ia et a i (1977, p. 146), the outer stereozone is completely abraded in longitudinal section so that the structure o f the dissepimentarium cannot be observed. In Temnophyllum mosaicusum K o n g , 1978 from the G ive­tian o f the Guizhou Province described by K o n g <&

H u a n g (1978, p. 96), the major septa are strongly fusi­form. There are also some affinities between T. majus and T. waltheri Y o h , 1937 from the Givetian o f the Guangxi Province in China, especially with one o f the paratypes figured by Y o h (1937, pi. 7, fig. 3). However in the holotype from You (1937, pi. 7, fig. 1), the peripheral stereozone is not perfectly continuous in transverse section and does not appear in longitudinal section. Moreover, the other paratype illustrated by Y o h (1937, pi. 7, fig. 2) probably belongs to the genus Spinophyllum as it was already suggested by B ir e n h e id e <& L ia o (1985, p. 243).

G e o g r a p h ic a n d s t r a t ig r a p h ic o c c u r r e n c e

The material sampled by the author comes mainly from the lower part o f the Mont d ’Haurs Formation at Nismes, Beauraing, Wellin, Ave-et-Auffe and Han-sur-Lesse, on the south side o f the Dinant Synclinorium. However, some specimens have been collected somewhat higher in several o f these localities.

Outside Belgium and Givet in France, Temnophyllum majus occurs only in the Givetian Schwelm Formation from the Sauerland in Germany.

Tem nophyllum im perfectum n. sp.Plate 2, Figures 1-10

v p. 1977 Temnophyllum latum- T sien , fig. 4n (nonfigs. 4o , p = Temnophyllum majus W a l t h e r . 1929).

(v non 1929 Temnophyllum latum- W a l t h e r , p. 123. fig. 14).

D erivado nominisFrom imperfectus (latin) = incomplete, referring to the incomplete stereozone o f the species.

HolotypeIRScNB a 11727 (= PI. 2, Figs. I, 2). Specimen Olloy- sur-Viroin MC-64-C824 collected by C o e n - A u b e r t in 1999, 6 m above the lower part o f the Mont d 'H aurs Formation.

Locus typicusSmall excavation (Olloy-sur-Viroin MC-64; Fig. 4) lo­cated in the Bois des Abannets, 1250 m to the northeast o f

14 Marie COEN-AUBERT

Nismes. Map sheet O lloy-sur-Viroin IGNB 58/5, Lam­bert coordinates: x= 164.125 and y= 85.125, south side o f the Dinant Synclinorium, Belgium.

Stratum typicumBase o f the upper part o f the Mont d ’Haurs Formation, middle part o f the Givetian.

Material and localitiesSeventy-two specimens with 111 thin sections. Personal sampling: Wellin MC-1988-6-A705, A799, B111G, B il l i , BÍ UL, BÍ 18, B 120, B 176 and B243; Wellin MC-1986-4- A510C, A510G, A511, A526C, A531E, A535N, A544D, A548 and A567; Beauraing MC-1975-3-Z685, Z686 and Z704; llan-sur-Lesse MC-1975-15-23, 83. 91, 94. 105, 108, 109, 110. 119 and 121: Olloy-sur-Viroin MC-64-C823, C824, C825, ('829, C'830, C831, C832. C833 and C835; Olloy-sur- Viroin MC-65-C897 and C898; G o z c e MC-47-C5I7 and C518; Nalinnes MC-1975-2-X38; Tamines MC-1983-2-Z5526, Z55213, Z55224, Z55227 and Z55229; Leuze-borehole 125E/ 298 at 1242 m. Old collection from the Institut royal des Sciences naturelles de Belgique: Agimont (Mont d’Haurs at Givet)-Gi-28386, 28432, 28437, 28468, 28470, 28493, 28496, 28520, 28526, 28528, 28570, 28596, 28610, 28615, 28631, 28643, 29021. 29044 and 29072; Olloy-sur-Viroin 12-Gid- 11985 and 12404.

D ia g n o s is

A species o f Temnophyllum with 56 to 68 septa at a diam eter o f 12 mm to 20 mm. Incomplete and variable stereozone developed in the outer part o f the dissepim en­tarium. Possible occurrence o f a few carinae.

D e s c r ip t io n

The material consists o f conical, cylindrical and some­times ceratoid coralla which are frequently fragmentary; their height varies between 1.5 cm and 5 cm. Growth lines and longitudinal ribs have been observed in a few speci­mens; the calice is deep rather rarely. A corallum shows lateral offsets whereas another one is affected by rejuve­nescence. The outer wall is not often well preserved. Some coralla are encrusted by thin laminar stromatopor­oids or even by auloporids, alveolitids, thamnoporids and fistuliporids.

The septa are non-carinate and typically dilated in the dissepimentarium. However, a few spinose or knobbly carinae occur occasionally; they are locally stronger in rare specimens. The septa are thinner, attenuated or rather thick in the tabularium. Sometimes, they are character­ized by a dark median line in the dissepimentarium. A peripheral stereozone, which is never continuous, is more or less developed against the wall and between the septa in the outer part o f the dissepimentarium. Occasionally also, a stereoplasmic thickening affects a layer o f dissepi­ments.

The m ajor septa reach the axis o f the corallum or leave a small open space in the centre o f the tabularium which is more extensive in a few specimens. The inner ends o f the m ajor septa may be rhopaloid, curved, forked or fused axially to form a pseudofossula. The minor septa traverse the entire dissepimentarium or even enter into the tabu­

larium where they are sometimes contratingent. They are rarely shorter or discontinuous at their axial ends.

The dissepim entarium consists o f 7 to 14 or even o f 5 to 17 rows o f small globose dissepim ents which are in horizontal layers at the periphery and inclined in its inner part. Some spots o f coarse trabeculae occur in the dis­sepimentarium whereas an outer stereozone is only pre­sent in a few specimens. The tabulae are incomplete and intersecting laterally; their axial parts are rarely convex or flat-topped.

There are 50 to 72 septa per corallum. The diameter o f the corallum ranges from 10 mm to 25 mm. The width o f the tabularium varies commonly between 6 mm and 10 mm and more generally between 4.5 mm and 13 mm.

D is c u s s io n

Temnophyllum imperfectum is very close to T. m ajus, especially by its quantitative data, that is to say by nearly sim ilar num ber o f septa, diam eter o f the corallum and o f the tabularium. In fact, the former species differs mainly from the latter in having an incomplete stereozone that does not invest all the outer part o f the dissepimentarium. That is why one o f the coralla figured by T s ie n (1977) as T. latum is referred herein to T. imperfectum. For the same reason, it is also possible that the sample from the Lower Givetian o f the Eifel Hills figured by S c h r ö d e r &

S a le rn o (2001, pi. 2, fig. 19) as T. sp. cf. latum belongs to the new species; but, it is partially abraded in trans­verse section and its diam eter is rather small.

Some carinae are present in a few specimens o f T. imperfectum. These specimens look more or less like the Givetian material from the lower part o f the Blacourt Formation in the Boulonnais, France, that was assigned to T. longiseptatum ( L ü t t e , 1984) by R o h a r t (1988, p. 275). However, the latter species was described in the genus Charactophyllum S i m p s o n , 1900 by L ü t t e

(1984, p. 184) and transferred to the genus Spinophyllum by L ü t t e & O e k e n t o r p (1988, p. 34), B ir e n h e id e &

L ü t t e (1990, p. 6) and W r z o l e k & W a c h (1994, p. 53). Indeed, the German sampling o f S. longiseptatum which comes from the Givetian Kerpen Formation o f the Eifel Hills, is characterized by more numerous and stron­ger carinae than Temnophyllum imperfectum.

T. neospongiosum ( I v a n i a , 1957) from the Givetian o f the Kuznetsk Basin in Russia is also a species that was ascribed to the genus Charactophyllum by I v a n ia (1957, p. 63 and 1965, p. 101). Though it has several features in common with Temnophyllum imperfectum , it is once more distinguished from the new taxon by its better developed carination. As for T. hreviseptatum F a n ,

1988 in H e & F a n (1988, p. 183) from the Devonian o f the Sichuan Province in China, it resem bles T. imperfec­tum by its incomplete and variable stereozone, but is separated from it by rather small coralla.

G e o g r a p h ic a n d s t r a t ig r a p h ic o c c u r r e n c e

The species is only known in the middle part o f the Givetian from Belgium and Givet in France. The material sampled by the author comes mainly from the south side

Mont d 'Haurs Formation 15

o f the Dinant Synclinorium where it has been found in the lower part o f the Mont d ’Haurs Formation at Ave-et- Auffe, Wellin, Beauraing, Han-sur-Lesse and Nismes and also somewhat higher in the three latter localities. Some specimens have been collected:

in the lower part o f the Mont d 'H aurs Formation at Cour-sur-Heure, on the north side o f the Dinant Syn­clinorium;

- in the lower part o f the Névremont Formation at Ger­pinnes, on the north side o f the Dinant Synclinorium and at Aisemont, on the south side o f the Namur Synclinorium;

- in the middle part o f the Alvaux Formation at Leuze, on the north side o f the Namur Synclinorium.The corallum figured by T s ie n (1977) comes from the

Mont d ’Haurs Formation at Givet.

Genus Spinophyllum W e d e k i n d , 1922

= Truncicarinulum Yu & K u a n g , 1982 = Charisphyllum O l iv e r & S o r a u f , 1988 = Spongielasma C a o , 1983 in C a o et al. (1983)

Type speciesBy monotypy, Campophyllum spongiosum S c h l ü t e r , 18 8 9 .

D ia g n o s is

Solitary rugose corals. Septa o f two orders, highly and ir­regularly carínate in the dissepimentarium with yardarm and zigzag carinae. Septa more or les dilated in the disse­pimentarium and thin in the tabularium. Major septa rea­ching the axis o f the corallum or leaving an open space in the centre o f the tabularium. Minor septa traversing the entire dissepimentarium. Dissepimentarium composed o f several rows o f globose dissepiments arranged in horizon­tal layers in its outer part and inclined towards the axis o f the corallum in its inner part. Tabulae incomplete or compound.

Spinophyllum spongiosum ( S c h l ü t e r , 1 8 8 9 )Plate 2, Figures 11, 12, Plate 3, Figures 3-7

v * 1889 Campophyllum spongiosum Schliit.- Schlü ter ,p. 46.

1922 Spinophyllum spongiosum Schlüter- W ed ek in d , fig. 2.

non 1951 Charactophyllum spongiosum (Schlüter)- S o sh k i- n a , p. 71, pi. 12.

non 1952 Charactophyllum spongiosum (Schlüter)- So sh k i- n a , p. 91, pi. 28.

non 1960 Charactophyllum spongiosum (Schlüter, 1889)- Z h eltonogova & Ivan ia , p. 402 , pi. D -53 , fig. 4, pi. D -54, fig. 1.

non 1963 Charactophyllum spongiosum (Schlüter)- Sm e- lo v sk a y a , p. 205 , pi. 42 , fig. 5, pi. 43 , figs. 5-7.

non 1965 CharactophvUum spongiosum (Schlüter)- Iv an ia , p. 102, pi. 54, figs. 245 , 246 , pi. 55, figs. 249, 250.

v 1967 Cylindrophyllum (Cylindrophyllum) spongiosum (Schlüter 1889)- P ick ett , p. 55.

1978 Charactophyllum spongiosum (Schlüter 1889)- B i­r e n h e id e , p. 85, pi. 12 fig . I.

1981 Spinophyllum spongiosum (Schlüter)- H il l , fig. 172', 3.

non 1987 Spinophyllum spongiosum (Schlüter)- C a o & OUYANG, p . 172, p i. 30, fig . 4.

v p. 1990 Spinophyllum spongiosum (Schlüter 1889)- B ir e n ­h e id e & L ü t t e , p . 4, pi. 1, figs. 4,6-8, pi. 2, figs. 10- 14, pi. 3, 15-20 (non pi. 1, figs. 1-3, 5, 9).

v 1994 Spinophvllum spongiosum (Schlüter)- W r z o l e k & W a c h , p i. 2, fig. 11.

LectotypePI. 3, fig 15 in B ir e n h e id e & L ü t t e ( 1990), pi. 2, fig. 11 in W r z o l e k & W a c h (1994) and pi. 2, figs. 11, 12 figured herein, chosen by B ir e n h e id e & L ü t t e (1990, p. 4). Specimen Nr 174a o f the S c h l ü t e r collection stored in the Paläontologisches Institut from the Univer­sity o f Bonn in Germany. Givetian Büchel Formation o f the disused Büchel quarry near Herrenstrunden and Ber- gisch Gladbach in the Bergisches Land, Germany.

Material and localitiesTen specimens with 15 thin sections. Personal sampling: Wel­lin MC-1988-6-A704, B150 and B241; Wellin MC-1974-95- Z841, Z847 and Z850; Tamines MC-1983-2-Z55212. Old col­lection from the Institut royal des Sciences naturelles de Belgi­que: Agimont (Mont d'Haurs at Givet)-Gi-28309, 28310 and 28841.

D ia g n o s is

A species o f Spinophyllum with 60 to 70 septa at a diam eter 13 mm to 21 mm. Yardarm and zigzag carinae rather strong and numerous. No stereoplasmic thickening between the septa in the adult stage.

D e s c r ip t io n

The material consists o f fragments o f conical, ceratoid and cylindrical coralla whose height varies between 0.8 cm and 5.5 cm. Longitudinal ribs are occasionally present. The outer wall is not well preserved though it may be rather thick. It is locally encrusted by thin laminar stromatoporoids or auloporids in a few specimens.

The septa bear yardarm, zigzag, spinose and knobbly carinae which are more or less numerous and strong. They are dilated in the dissepimentarium and become thinner or sometimes less thick in the tabularium. In some specimens, the septa are locally slender or discontinuous at the periphery. A rare deposit o f stereoplasma is present within the dissepimentarium o f a few coralla.

The m ajor septa reach the axis o f the corallum or leave an open space in the centre o f the tabularium. O ccasion­ally, their inner ends are discontinuous, fusing axially or forming pseudofossulae; small spinose carinae occur in the tabularium o f one sample. The m inor septa traverse the entire dissepimentarium or enter the tabularium where they may be contratingent; they are rarely discontinuous at their axial ends.

The dissepimentarium consists o f 8 to 14 rows o f small dissepiments which are in horizontal layers at the periph-

16 Marie COEN-AUBERT

ery and inclined in its inner part. Coarse trabeculae, which are frequently isolated and only locally contiguous, extend across the entire dissepimentarium. The tabulae are incomplete and intersecting laterally; their axial parts are sometimes horizontal or flat-topped.

There are 62 to 76 septa per corallum. The diameter of the corallum ranges from 11.5 mm to 25 mm whereas the width o f the tabularium varies between 5 mm and 10.5 mm.

D is c u s s io n

As Spinophyllum spongiosum is the type species o f Spi­nophyllum, it is a pity that B i r e n h e id e & L ü t t e (1990) chose as lectotype a specimen whose transverse section belongs to a juvenile stage (PI. 2, fig. 11). In this trans­verse section, one can see more or less strong zigzag carinae, but also stereoplasmic thickenings in some parts o f the dissepimentarium. However, these stereoplasmic thickenings do not occur in the German adult coralla o f S. spongiosum figured by W e d e k in d (1922), B i r e n h e id e

(1978) and H il l (1981). Among the abundant material illustrated by B ir e n h e id e & L ü t t e (1990), some stereo­plasmic thickenings are present, but I agree with W r z o ­

l e k & W a c h (1994, p. 53) that such specimens must be assigned to S. longiseptatum.

Differences in the carination mainly justify the exclu­sion o f the Russian and Chinese references from the synonymy o f S. spongiosum. These carinae are poorly developed in the Givetian sample from the Qinling M ountains investigated by C a o & O u y a n g (1987) as well as in part o f the Givetian material from the Urals figured by S o s h k in a (1951, pi- 12, fig. 1 and 1952). The other corallum illustrated in thin sections by S o s h k in a

(1951, pi. 12, fig. 3 and 1952) is much larger than 5. spongiosum and shows typically dilated septa with nu­merous yardarm carinae. Some strong carinae occur only at the periphery o f the specimens collected by S m e l o v s -

k a y a (1963) in the Givetian from the Tarbagatai ridge. Finally, weak carinae are confined to the external part o f rather thick septa in the sampling o f Z h e l t o n o g o v a &

Iv a n ia (1960) and Iv a n ia (1965) coming from the G ive­tian o f the Kuznetsk Basin.

S. spongiosum has several features in common with the three paratypes o f S. altevogti ( O l iv e r & S o r a u f , 1988) from the Givetian o f Asturias in Spain figured by O l iv e r

& S o r a u f (1988, figs. 6, 7). However the holotype o f S. altevogti is different as it is characterized by numerous yardarm carinae throughout the dissepimentarium and by the occurrence o f third order septa which are short or discontinuous. Indeed, this taxon was described as Helio- phyllum tricyclicum C h e n g , 1969 in an unpublished the­sis by C h e n g (1969). It must also be mentioned that Spinophyllum altevogti is the type species o f the genus Charisphyllum O l iv e r & S o r a u f , 1988 and that the two transverse sections o f its holotype are reillustrated herein (PI. 3, figs. 1,2).

Spinophyllum columellum ( C a o , 1983 in C a o et al., 1983, p. 74) from the Devonian o f Northwest China and type species o f Spongielasma C a o , 1983 in C a o et al. (1983) is also very close to Spinophyllum spongiosum

though it has slightly fewer septa and rather thick septa like the paratypes o f S. altevogti.

S. aiense ( S o s h k in a , 1949) from the Givetian o f the Urals in Russia is another taxon that resembles S. spongio­sum. However, it is distinguished from it by weaker cari­nae and by major septa reaching systematically the centre o f the tabularium. Mostly zigzag carinae characterize also:- S. puanense ( K o n g , 1978) described in the subgenus

Cyathophyllum (Peripaedium) E h r e n b e r g , 1834 by K o n g & H u a n g (1978, p. 103) and collected near the Lower-M iddle Devonian boundary in the Guizhou Province, China;

- Spinophyllum poshiense ( F o n t a i n e , 1966) from the Middle Devonian o f the Yunnan Province in China which was also ascribed to Peripaedium by F o n t a in e

(1966, p. 55) and which differs additionally from Spi­nophyllum spongiosum by more septa and dissepiments;

- S. trochoides ( H il l , 1942) from the Givetian o f North Queensland in Australia which has slightly fewer septa than S. spongiosum and which was transferred to Spi­nophyllum by Z i ie n & J e l l (1996, p. 76).As for S. zhongguoense (He, 1978) from the Givetian o f

the Sichuan Province in China and assigned to Spinophyl­lum by B i r e n h e id e & L ia o (1985, p. 245), it is mainly separated from S. spongiosum by less dilated septa.

G e o g r a p h ic a n d s t r a t ig r a p h ic o c c u r r e n c e The material sampled by the author comes mainly from the top o f the Terres d ’Haurs Formation and from the lower part o f the Mont d ’Haurs Formation at Ave-et- Auffe and Resteigne, on the south side o f the Dinant Synclinorium. One specimen has also been collected in the lower part o f the Névrem ont Formation at Aisemont, on the south side o f the N am ur Synclinorium.

Outside Belgium and Givet in France, Spinophyllum spongiosum occurs in the middle part o f the Givetian from several areas in Germany, that is to say in the Kerpen Formation from the Eifel Hills, in the Büchel Formation from the Bergisches Land and in the Schwelm Formation from the Sauerland.

Spinophyllum blacourti (R on A R T, 1988)Plate 3, Figures 8-14

* 1988 Truncicarinulum blacourti nov. sp. - R o h a r t ,

p. 277, pi. 35, figs. 3, 4.

HolotypePI. 35, fig. 3 in R o h a r t (1988). Specimen GFCL 4556 stored in the Department o f Geology from the Faculté Libre des Sciences at Lille, France. Banc Noir quarry at Ferques, Boulonnais, France. Unit c o f the Griset M em­ber, Blacourt Formation, middle part o f the Givetian.

Materia! and localitiesThirty-nine specimens with 58 thin sections. Personal sam­pling: Wellin MC-1988-6-A724, A756, B3, B7, B ll IB, BÍ 11C, BÍ 110, B 115, B 168, B173 and B242; Wellin MC- 1986-4-A535B, A535H and A544A; Beauraing MC-1975-3-

Mont d ’Haurs Formation 17

Z689; Gozée MC-47-C496 and C497; Nalinnes MC-1975-2-81; Famines MC-1983-2-Z5521, Z5527, Z5528, Z5529, Z55214, Z55216, Z55217, Z55223, Z55228 and Z560. Old collection from the Institut royal des Sciences naturelles de Belgique: Agimont (Mont d'Haurs at Givet)-Gi-28379, 28555, 28556, 28574, 28617, 28702, 28850, 28858, 28940 and 29047; Ol­loy-sur-Viroin 12-Gid-l 1924.

D ia g n o s is

A species o f Spinophyllum with 56 to 68 septa at a diameter o f 15 mm to 25 mm. Septa slightly carínate with rare yardarm carinae.

D e s c r ip t io n

The material consists o f conical, trochoid, ceratoid and cylindrical coralla which are often fragmentary and show frequent longitudinal ribs. Their height varies between 1 cm and 5.5 cm, but may reach 7 cm or even 9.5 cm. Rare laterals offsets have been observed. The outer wall is not very well preserved and is encrusted by thin laminar stromatoporoids in only a few specimens.

The septa are non-carinate or bear in the outer part o f the dissepimentarium some spinose, knobbly or yardarm carinae which are normally weak and sometimes stron­ger. The septa are more or less dilated in the dissepim en­tarium and become thinner or less thick in the tabularium. Occasionally, they are dilated throughout their length, but they may also be slender in the dissepimentarium or at least at the periphery o f the corallum. In several speci­mens, there is locally a stereoplasmic thickening against the outer wall or on a layer o f dissepiments rather near it.

The m ajor septa reach the axis o f the corallum or leave an open space in the centre o f the tabularium. Their inner ends are sometimes rhopaloid, discontinuous, curved or fusing axially to form pseudofossulae or possibly a plane o f bilateral symmetry. The m inor septa traverse the entire dissepimentarium or enter occasionally into the tabular­ium where they may be contratingent; they are rarely shorter or discontinuous at their axial ends.

The dissepimentarium consists o f 6 to 13 or even 2 to 14 rows o f small dissepiments which are normally in horizon­tal layers at the periphery and inclined in its inner part. The tabulae are incomplete and intersecting laterally; in a few coralla, they are locally horizontal or concave.

There are 52 to 76 septa per corallum. The diam eter o f the corallum ranges from 12 mm to 22 mm or even from9.5 mm to 26 mm. The width o f the tabularium varies commonly between 6 mm and 10 mm and more generally between 4.7 mm and 11 mm.

D is c u s s io n

The investigated material is very similar to that o f R o ­h a r t (1988) though it is often characterized by slightly sm aller coralla. The specimen from the Givetian Kerpen Formation in the Eifel Hills described by L ü t t e (1984, p. 186) as Charactophyllum sp. was assigned by R o h a r t

(1988) to Spinophyllum blacourti; however, it is much more carínate and seems to be closer to S. spongiosum.

Two species from the same lithostratigraphic unit and area in Germany are more or less related to S. blacourti.

Firstly, 5. arduum (L ü t t e , 1 9 8 5 ) is another taxon o f Spinophyllum with poorly developed carinae that differs from S. blacourti by thinner septa. It was ascribed to the colonial genus Cyathophyllum G o l d f u s s , 1 8 2 6 by L ü t t e ( 1 9 8 5 , p. 5 4 4 ) and transferred to Spinophyllum by Ro­h a r t ( 1 9 8 8 , p. 2 8 4 ) who found it together with S. bla­courti in the Boulonnais. Secondly, the holotype o f M ic- tophyllum schlueteri B ir e n h e id e & L ü t t e , 1 9 9 0 resem­bles Spinophyllum blacourti, but the other coralla figured by the authors are different in having m inor septa o f highly variable length. It must also be mentioned that Charactophyllum eguchii M a , 1 9 5 6 from the Middle Devonian o f the Eifel Hills introduced by M a ( 1 9 5 6 , p. 43) is mainly separated from Spinophyllum blacourti by septa only locally dilated in the dissepimentarium.

S. spongiosum is easily distinguished from S. blacourti by its stronger and more numerous carinae which are frequently o f the yardarm type; moreover, it has a few more septa. By the rather scarcity o f its carination, S. blacourti occupies a marginal position in the genus Spino­phyllum. This is also the case for Keriophyllum temenio- phylloides W a n g , 1948 from the Middle Devonian o f the Yunnan Province in China which was referred to the subgenus Temnophyllum (Temnocarinia) Y u & L ia o ,

1 97 8 by K o n g & H u a n g (1 9 7 8 , p. 1 00). However, Tem­nocarinia is a nomen nudum that was redescribed by Y u & K u a n g (1 9 8 2 , p. 2 5 3 ) as Truncicarinulum Y u & K u a n g ,

1 982 synonym o f Spinophyllum according among others to M c L e a n (1 9 9 3 , p. 110) and W r z o l e k & W a c h (1 9 9 4 , p. 5 3 ). As for S. temeniophylloides, it differs from S. blacourti by smaller size and septal number.

G e o g r a p h ic a n d s t r a t ig r a p h ic o c c u r r e n c e The material sampled by the author at Beauraing, W ellin and Ave-et-Auffe, on the south side o f the Dinant Syn­clinorium, comes from the lower part o f the Mont d ’Haurs Formation. Moreover, several specimens have been collected in the lower part o f the N évrem ont For­mation at Aisemont, on the south side o f the Namur Synclinorium and at Gerpinnes, on the north side o f the Dinant Synclinorium. In the latter area, a few coralla have also been found close to the top o f the Terres d 'H aurs Formation at Cour-sur-Heure.

Outside Belgium and Givet in France, Spinophyllum blacourti occurs only in the lower part o f the Griset M ember from the Givetian Blacourt Formation in the Boulonnais, France.

A c k n o w l e d g e m e n t s

E. S c h i n d l e r (Frankfurt am M ain), M. S a n d e r (B onn) and K. O e k e n - t o r p (M ünster) kindly let m e study type specim ens in their care. M. C o e n (Louvain-la-N euve) and V irginie D u m o u l i n (B ruxelles) show ed m e the N ism es exposures. W .A . O l i v e r , Jr. (W ashington,D .C .), D enise B r i c e (Lille) and B . M i s t i a e n (L ille) review ed and helpfully com m ented a previous version o f the paper. Thin sections w ere prepared by R. C r e m e r s (IR ScN B) and by the Laboratory o fE. P o t y (Liège). Photos w ere prin ted by W. M i s e u r (IR ScNB). T o all these colleagues, my deep gratitude.

18 Marie COEN-AUBERT

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F r e c h , F., 1886. Die Cyathophylliden und Zaphrentiden des deutschen Mitteldcvon. Palaeontologische Abhandlungen, 3 (3): 1-120.G a l l e , A., 1985. Biostratigraphy and rugose corals of Mora­vian Devonian (Czechoslovakia). Newsletters on Stratigraphy, 14: 48-68.G a l l e , A., 1994. Rugose corals of the Acanthopyge Limestone of Koneprusy (Middle Devonian, Barrandian, Czech Republic). Vestnik Ceskeho geologickeho uslavu, 69: 41-58.G o l d f u s s , A., 1826. Petrefacta Germaniae 1: 1-76. Amz & Comp.. Düsseldorf.G ü r ic h , G., 1896. Das Paleozoicum im Polnischen Mittelge­birge. Verhandlungen der Russisch-kaiserlichen Mineralo­gischen Gesellschaft zu St. Petersburg, (2), 32: 1-539.H e , Y.X., 1978. Subclass Rugosa. In: Chengdu Institute of Geology and Mineral Resources (Editor), Atlas of fossils of Southwest China. Sichuan Volume. Part I. From Sinian to Devonian. Geological Publishing House, Beijing, pp. 98-178.He, Y.X. & F a n , Y.N., 1988. Rugosa. In: Hou, H.F. (Editor), Devonian stratigraphy, paleontology and sedimentary facies of Longmenshan, Sichuan. Geological Publishing House, Beijing, pp 165-195.H il l , D., 1939. The Devonian rugose corals of Lilydale and Loyola, Victoria. Proceedings o f the Royal Society o f Victoria, new series, 51: 219-256.H il l , D., 1942. The Middle Devonian rugose corals of Queens­land, III. Burdekin Downs, Fanning R„ and Reid Gap, North Queensland. Proceedings o f the Royal Society o f Queensland, 53: 229-268.H il l , D., 1981. Part F, Coelenterata, Supplement I, Rugosa and Tabulata, 2 vols. In: T e ic h e r t , C. (Editor), Treatise on Inverte­brate Paleontology. The Geological Society of America, Inc. and The University of Kansas, Boulder, Colorado and Law­rence, Kansas, 762 pp.Iv a n ia , V.A., 1957. Devonskie korally der. Belyi Kamen na r. Kara-Chumysh (yugo-zapadnyi Kuzbass). Uchenye Zapiski Tomskogo Gosudarstvennogo Universiteta, 28: 55-73 (in Rus­sian).Iv a n ia , V.A., 1965. Devonskie korally Sayano-Altayskoy gor- noy oblasti. Izd. Tomskogo Universiteta, Tomsk, 398 pp (in Russian).J ia , H.Z., Xu, S.Y., K u a n g , G.D., Z h a n g , B.F., Z h u o , Z.B. & Wu, J.S., 1977. Anthozoa. In: Hubei Provincial Geological Science Research Institute (Editor), Atlas of the paleontology of the South Central Regions, part 2, Late Paleozoic. Geological Publishing House, Beijing, pp. 109-270.J o s e p h , J. & T s ie n , H.H., 1975. Calcaires mésodévoniens et leurs faunes de Tétracoralliaires en Haute Vallée d'Ossau (Py- rénées-Atlantiques). Bulletin de la Société d'Histoire naturelle de Toulouse, 111: 179-203.K e t t n e r o v a , M., 1932. Paleontologicke studie z celechovic- keho devonu, Cast IV: Rugosa. Prace geologicko-palaeontolo- gickeho ustavu Karlovy university v Praze, 1932: 1-97.K o n g , L. & H u a n g , Y.M., 1978. Tetracoralla. In: Guizhou Stratigraphy and Palaeontology Work Team (Editor), Palaeon­tological Atlas of Southwest China. Guizhou Volume. Part 1, Cambrian-Devonian. Geological Publishing House, Beijing, pp. 35-161.

Mont d ’Haurs Formation 19

K r ä m e r , T.J., 1982. Cerioide Rugosa aus dem Devon der Bergisch Gladbach-Paffratlier Mulde (Rheinisches Schieferge­birge). Neues Jahrbuch Jur Geologie und Paläontologie, Mo­natshefte, 1982 (11): 648-666.L a n g , W .D ., S m ith , S. & T h om a s , H.D., 1940. Index of Pa­laeozoic Coral genera. British Museum (Natural History), Lon­don, 231 pp.L ü tte , B .P ., 1984. Rugose Korallen aus dem Mitteldevon (Givetium) der Sötenicher Mulde (Rheinisches Schieferge­birge, Nord-Eifel). Münstersche Forschungen zur Geologie und Paläontologie, 61: 175-243.

L ü t t e , B.P., 1985. Cyathophyllum (Cyathophyllum) arduum n. sp. (Rugosa) aus der Sötenicher Mulde (Rheinisches Schiefer­gebirge, Nord-Eifel). Neues Jahrbuch Jur Geologie und Pa­läontologie, Monatshefte, 1985 (9): 542-552.L ü t t e , B.P. & O e k e n t o r p , K., 1988. Die stratigraphische Verteilung der solitären Rugosa im Givetium (Mittel-Devon) der Eifel (Rhenisches Schiefergebirge). Newsletter on Strati­graphy, 20: 29-42.Ma, T. Y.H., 1956. A reinvestigation of climate and the relative positions of continents during the Devonian. Research on the Past Climate and Continental Drift, 9: 1-116.M a y , A ., 1993. Korallen aus dem höheren Eifelium und unteren Givetium (Devon) des nordwestlichen Sauerlandes (Rhei­nisches Schiefergebirge). Teil II: Rugose Korallen, Chaetetiden und spezielle Themen. Palaeontographica, A, 228: 1-103.

M cL e a n , R .A ., 1993. The Devonian rugose coral Family Char- actophyllidae P e d d e r . Courier Forschungsinstitut Sencken- berg, 164: 109-118.M id d le to n , G .V ., 1959. Devonian tetracorals from South De­vonshire, England. Journal o f Paleontology, 33: 138-160.

O l iv e r , W .A . & S o r a u f , J.E., 1988. Hedophyllum H a l l and Charisphyllum n. gen. (Devonian rugose corals) of the Canta­brian Mountains (N W Spain). Trabajos de Geología, Universi­dad de Oviedo, 17: 3-17.

P ickett , J., 1967. Untersuchungen zur Familie Phillipsastreidae (Zoantharia rugosa). Senckenbergiana lethaea, 48 (1): 1-89.

R o h a r t , J.C., 1988. Rugueux givetiens et frasniens de Ferques (Boulonnais-France). In: B r ic e , D. (Editeur), Le Dévonien de Ferques. Bas-Boulonnais (N. France). Biostratigraphie du Pa- lèozoïque, 7: 231-297.S c h l ü t e r , C., 1889. Anthozoen des rheinischen Mittel-Devon. Abhandlungen zur geologischen Specialkarte von Preussen und der Thüringischen Staaten, 8 (4): 1-207.S c h r ö d e r , S. & S a l e r n o , C., 2001. Korallenfauna und Fazies givetischer Kalksteinabfolgen (Cürten-/Dreimühlen-Forma- tion) der Dollendorfer Mulde (Devon, Rheinisches Schieferge­birge/Eifel). Senckenbergiana lethaea, 81: 111-133.S im p s o n , G. B., 1900. Preliminary descriptions of new genera of Paleozoic rugose corals. Bulletin o f the New York State Mu­seum: 8 (37): 199-222.S m e l o v s k a y a , M.M., 1963. Rugozy srednego Devona. In: B o g d a n o v , A .A . (Editor), Stratigrafíya i fauna Paleozoyskikh otlozheniy Khrebta Tarbagatai (Ordovik. Silur, Devon, Nijniy Karbon). Gostgeoltekhizdat, Moskva, pp. 195-206 (in Russian).S o s h k in a , E.D., 1949. Devonskie korally Rugosa Urala. Trudy Paleontologicheskogo Instituía, 15 (4): 1-160 (in Russian).S o s h k in a , E.D., 1951. Pozdnedevonskie korally Rugosa, ikh sistematika i evolyutsiya. Trudy Paleontologicheskogo Institu­ía, 34: 1-122 (in Russian).

S o s h k in a , E.D., 1952. Opredelitel’ devonskikh chetyrekhlu- chevykh korallov. Trudy Paleontologicheskogo Instituto, 39: 1-127 (in Russian).T s ie n , H.H., 1975. Introduction to the Devonian Reef develop­ment in Belgium. Livret-Guide, Excursion C (Nord de la France et de la Belgique), 2e Symposium International sur les Coraux et Récifs coralliens fossiles, Paris 1975. Bruxelles, pp. 3-43.T s ie n , H.H., 1977. The sequence and distribution of Frasnian rugose corals fauna in Belgium. Mémoires du Bureau de re­cherches géologiques et minières, 89: 203-220.T s ie n , H.H., 1978. Rugosa massifs du Dévonien de la Belgique. Mémoires de l'Institut Géologique de l'Université de Louvain, 29: 197-229.V o n S c h o u p p é , A., 1951. Kritische Betrachtungen und Revi­sion des Genusbegriffes Entelophyllum WDK. nebst einigen Bemerkungen zu W e d e k in d s “ Kypophyllidae” und “ Kodono- phyllidae” . Sitzungsberichte der Österreichische Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Klasse, I, 160: 243-256.V o n S c h o u p p é , A., 1954. Korallen und Stromatoporen aus dem ef der Kamischen Alpen. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 99: 379-450.W a l t h e r , C., 1929. Untersuchungen über die Mitteldevon- Oberdevongrenze. Zeitschrift der Deutschen Geologischen Ge­sellschaft, 80: 97-152.W a n g , H.C., 1948. The Middle Devonian rugose corals of Eastern Yunnan. Contributions from the Geological Institute, National University o f Peking, 33: 1-45.W e d d ig e , K. (Editor), 1996. Devon-Korrelationstabelle. Senck­enbergiana lethaea, 76: 267-286.W e d d ig e , K. (Editor), 1998. Devon-Korrelationstabelle. Ergän­zungen 1997. Senckenbergiana lethaea, 77: 289-326.W e d d ig e , K. (Editor), 2000. Devonian Correlation Table. Sup­plements 2000. Senckenbergiana lethaea, 80: 691-726.W e d e k in d , R., 1922. Zur Kenntnis der Stringophyllen des oberen Mitteldevon. Sitzungsberichte der Gesellschaft zur Be­förderung der gesamten Naturwissenschaften zu Marburg, 1921 (1): 1-16.W e d e k in d , R., 1925. Das Mitteldevon der Eifel. Eine biostrati­graphische Studie. II. Teil. Materialien zur Kenntnis des mit­tleren Mitteldevon. Schriften der Gesellschaft zur Beförderung der gesamten Naturwissenschaften zur Marburg, 14 (4): 1-85.W r z o l e k , T., 1993. Rugose corals from the Devonian Kowala Formation of the Holy Cross Mountains. Acta Palaeontologica Polonica, 37: 217-254.W r z o l e k , T.& W a c h , P., 1994. Tetracoral genus Spinophyllum in the Devonian of the Holy Cross Mts, Poland. Geología, 12/ 13: 47-63.Y o h , S.S., 1937. Die Korallenfauna des Mitteldevons aus der Provinz Kwangsi, Südchina. Palaeontographica, A, 87: 45-76.Yu, C.M. & K u a n g , G.D., 1982. Late Middle Devonian rugose corals from Liujing, Heng Xian, Guangxi and their paleoeco- logical signifiance. Bulletin o f the Nanjing Institute o f Geology and Palaeontology, Academia Sinica, 4: 241-278.Z h e n , Y.Y. & J e l l , J.S, 1996. Middle Devonian rugose corals from the Fanning River Group, North Queensland, Australia. Palaeontographica, A, 2 42: 15-98.Z h e l t o n o g o v a , V.A. & Iv a n ia , V.A., 1960. Podklass Tetra- coralla (Rugosa). Tetrakorally. In: K h a l f in , L.L. (Editor),

20 Marie COEN-AUBERT

Biostratigrafiya paleozoya Sayano-Altayskoy gornoy oblasti. Trudy Sibirskogo nauchno-issledovatelskogo Instituto Geolo- gii, Geofiziki i Mineralnogo Syrya, 20: 368-408 (in Russian).

Marie C o e n -A u b e r t

Département de PaléontologieSection des Invertébrés fossilesInstitut royal des Sciences naturelles de Belgiquerue Vautier 29, B-1000 Bruxelles, BELGIUME-mail: Marie.Coen-Aubert@naturalsciences.be.

Typescript subm itted August 22, 2001 Revised typescript received October 25, 2001

Explanation of Plates

All specimens are figured at magnification x 3.

P l a t e 1

Temnophyllum latum W a l t h e r , 1929

Figs. 1,2 — Holotype. Thin sections SMF WDKD 6972 and 6971 stored in the Forschungsinstitut Senckenberg at Frankfurt amMain, Germany. Transverse and longitudinal sections.

Temnophyllum majus W a l t h e r , 1929

Figs. 3, 4 — Lectotype. Thin sections SMF WDKD 6996 and 6997 stored in the Forschungsinstitut Senckenberg at Frankfurtam Main, Germany. Transverse and longitudinal sections.

Figs. 5, 6 — Paralectotype. Thin sections SMF WDKD 6992 and 6994 stored in the Forschungsinstitut Senckenberg atFrankfurt am Main, Germany. Transverse and longitudinal sections.

Fig. 7 — Holotype o f Temnophyllum clavatum W a l t h e r , 1929. Thin section SMF W D K D 6977 stored in the Forschungs­institut Senckenberg at Frankfurt am Main, Germany. Transverse section.

Fig. 8 — IRScNB al 1722. Olloy-sur-Viroin MC-65-C867. Transverse section.Figs. 9, 10 IRScNB al 1723. Beauraing MC-1975-3-Z703. Transverse and longitudinal sections.Figs. 11, 12 — IRScNB al 1724. Wellin MC-1986-4-A510E. Transverse and longitudinal sections.Fig. 13 IRScNB al 1725. Wellin MC-1988-6-A723. Transverse section.

P l a t e 2

Temnophyllum imperfectum n. sp.

Figs. 1, 2 — Holotype. IRScNB al 1727. Olloy-sur-Viroin MC-64-C824. Transverse and longitudinal sections.Fig. 3 — Paratype. IRScNB al 1728. Olloy-sur-Viroin MC-65-C897. Transverse section.Figs. 4, 5 — Paratype. IRScNB al 1729. Han-sur-Lesse MC-1975-15-119. Transverse and longitudinal sections.Figs. 6, 7 — Paratype. IRScNB al 1730. Wellin MC-1986-4-A548. Transverse and longitudinal sections.Fig. 8 — Paratype. IRScNB a 11731. Beauraing MC-1975-3-Z704. Transverse section.Figs. 9, 10 — Paratype. IRScNB al 1732. Han-sur-Lesse MC-1975-15-110. Transverse and longitudinal sections.

Spinophyllum spongiosum (S c h l ü t e r , 1889)

Figs. II. 12 — Lectotype. Specimen Nr 174a of the S c h l ü t e r collection stored in the Paläontologisches Institut from the University of Bonn, Germany. Transverse and longitudinal sections.

Temnophyllum majus W a l t h e r , 1929

Figs. 13, 14 IRScNB al 1726. Wellin MC-1986-4-A532P. Transverse and longitudinal sections.

Mont d ’Haurs Formation 21

Figs. 1, 2

Figs. 3, 4 Figs. 5, 6 Fig. 7

Figs. 8, 9 Fig. 10 Figs. 11, 1 Figs. 13, I

P l a t e 3

Spinophyllum altevogti (O l iv e r & S o r a u f , 1988)

Holotype. Thin sections B2.28/1 and 2 stored in the Geologisch-Paläontologisches Institut und Museum from theUniversity of Münster, Germany. Transverse sections. Fig. 2 was figured by O liv er & S o r a u f (1988, fig. 5A).

Spinophyllum spongiosum (S ch lü ter , 1889)

— IRScNB al 1733. Wellin MC-1974-95-Z841. Transverse and longitudinal sections.— IRScNB al 1734. Wellin MC-1974-95-Z847. Transverse and longitudinal sections.— IRScNB a l l 735. Wellin MC-1988-6-B241. Transverse section.

Spinophyllum blacourti (R o h a r t , 1988)

— IRScNB aí 1736. Wellin MC-1988-6-B7. Transverse and longitudinal sections.— IRScNB al 1737. Tamines MC-1983-2-Z560. Transverse section.— IRScNB al 1738. Tamines MC-1983-2-Z55216. Transverse and longitudinal sections.

IRScNB al 1739. Beauraing MC-1975-3-Z689. Transverse and longitudinal sections.

2 2 Marie COEN-AUBERT

Pla te 1

Mont d ’Haurs Formation 23

P late 2

2 4 Marie COEN-AUBERT

P late 3